Based on everything the coaches have said, the SLB will be very different from last year. I would not expect Lee to play there at all. Remember, this is no longer the Rod M. 4-3 defense you probably played in high school.
Off-ball backers - (MLB and WLB) - Smith, LVE, Lee, Thomas, March, Covington, Benard
SLB/DPR - Anae, Gregory, Jelks, and Gifford (this is the spot he played at Nebraska, and had a good senior year playing that spot)
It is possible that Smith moves to SLB, which would weaken the off-ball spots, but if LVE is good to go, then Lee and Thomas are both capable starters and we have depth there.
I would not be surprised at all to see one more vet brought in to the SLB/DPR, especially if Gregory is not reinstated. Golden played that role for New York last year and did quite well and Matthews played that role for McCarthy in GB.