I reviewed some 2016 games.
He did play primarily as a DE, but it was normally more of 7-tech than 5-tech; although, he lined up in various places from 9-tech, 7-tech, 5-tech and DT. He didn't drop in coverage much, but they would do odd things like line him up at DT then drop him in coverage. He was in a heavy rotation and he was often not on the field. They also used #91 (Upshaw) and #94 (Tyron Jackson 6-4, 296) at LDE. The oddest thing is that he was often out of the lineup on 3rd and long with #91 or #94 at LDE.
They played a lot of wide splits with their DEs, occasionally in a wide-9 look and other times with a 7-9 look.
It's an interest defensive scheme. Their HC is the same guy that used the 320+ Red Bryant as the starting LDE for Seattle.