Parche
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You can best answer this by figuring out what they play against. There is more speed and off-the-snap acceleration at DE or OLB (usually). Tackles need to be quicker off the snap and it helps if they have long arms and quick feet. The long arms are so they can make contact with the pass rusher before the pass rusher makes contact with them. Hand placement, technique with the hands and upper-body strength are vital to deal with the combination speed-strength from NFL edge rushers. They need quick feet to deal with the moves and counter-moves from pass rushers who might start wide and dip or spin inside, or start in the body, then work outside. The quickness off the snap is a must at LT and really pretty necessary at RT, too. Otherwise the edge rusher can capture the corner off the snap and it's all over. What makes Tyron so good is he's solid off the snap and his hands are maybe the best in the league. Once he gets his hands on you, with his upper body strength, you're basically done. His feet are great, too.
At guard you work more in a small box, so you need to be stronger in the lower body (upper body strength helps too) but you can get by being a little slower off the snap because DT in the league are usually slower off the snap than an edge guy. At guard the leverage game is just as important as the speed game. If you can find a guard that can also move, that opens up all kinds of options and if you watch Zach Martin play you will see how.
One knock on Collins was poor hands in pass protection and average off-the-snap ability, which is why he projected better at guard. Free is much better with his hands but Free has lost a step in his lateral movement and get-off-the ball ability and that's why he seemed to get worse as the year went on this season. Collins also has slower feet, so he gets caught leaning with the upper body often. It's not usually a big deal at guard, but can be disastrous at tackle.
WOW, thanks. And yes, I understand what youre saying with Zach Martin. For example, quicker guards can excel and play pulls like him. For slower guards, surely youre not going many pulls at all.